Senior clerics had argued that the shrines should be kept open, and on Monday night crowds protesting against the closures forced their way into them.
Iran has temporarily freed thousands of additional prisoners, including political prisoners, in an attempt to reduce pressure on its prisons system as the country struggles to contain the spread of coronavirus.
"Things are much worse for the Iranian people, and we are convinced that will lead the Iranian people to rise up and change the behavior of the regime", Secretary of State Mike Pompeo said in early 2019. That has anxious health officials, who for weeks wanted Iran's Shiite clergy to close them.
At least 12 of those deaths have been of Iranian politicians and officials, both sitting and former, while 13 others have been infected and are now in quarantine and receiving treatment.
To encourage people to stay at home in Iran, Health Ministry spokesman Kianoush Jahanpour wrote on social media that the virus infects 50 Iranians on average every single hour and that 'one dies every 10 minutes'.
Underlining that urgency was the fatwa issued by Supreme Leader Ayatollah Khamenei, which prohibited "unnecessary" travel.
A key pilgrimage site in Iran's holy Shia city of Mashhad has been closed to prevent the spread of the novel coronavirus, a spokesman said on Sunday.
Among those temporarily freed was Nazanin Zaghari-Ratcliffe, an Iranian-British dual national long held on internationally criticized charges.
"Those who will be pardoned will not return to jail. nearly half of those security-related [political] prisoners will be pardoned as well".
"With the arrival of this batch of evacuees, the total number of quarantined people housed at the Army wellness facility in Jaisalmer rose to 484", an Army statement said.
It also locked down the Red Sea province that includes the resort town of Hurghada. It also halted all private sector work and public transportation as well.
A day later, he himself would be in quarantine from the virus.
In Oman, the sultanate announced anyone coming from overseas would be subject to quarantine. They tested negative for COVID-19 in the first test. * The IAF flight had also carried medical supplies which were given as a good-will gesture to China. Israel also called on citizens to stay home, closing parks, museums, libraries, beaches and other public areas.
In Syria, all sports clubs, movie theaters, concerts, theaters, as well as halls used for weddings or funerals were ordered closed in Damascus, and all restaurants and other shops around the country were ordered shut. Syria says it has no cases of the virus so far.
Dr Mandhari said a total of 18,000 confirmed cases of Covid-19 and more than 1,100 related deaths had been reported across the WHO's Eastern Mediterranean Region, which comprises 21 member states and the Palestinian territories. Gambrell reported from Dubai, United Arab Emirates. Associated Press writers Munir Ahmed in Islamabad, Pakistan and Sarah El Deeb in Beirut contributed.